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From the art book we know they were originally going to be the inhabitants of the Mongo river valley, worshipping, among other things, the annual return of the Gabbits to breed at Ma Spa.
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I know I read somewhere (and I think it was in the art book) that the muds in that picture were only put in because the grubbs had not yet been designed but they wanted to conceptualise the Mongo Valley. And of course, it has been said elsewhere of the Grubbs that they worshipped the Gabbit swarms.
I have no problem with the Grubbs. I think they are cute, funny, have an interesting ethos/collective personality and are distinctive enough for my tastes.