I think they are wachtowers because:
1. The machinary (I think) is to make electric energy for the great light of outside the wathtower, ( the mudokon run like a hamster to transform or store energy for the tower´s work)
2. in added the slig run up to a kind of gramophon... to give a alarm?
For me perhamps they are wath a great glukkon facility acces or better... the sligs are in the frontier with the wild territories of Oddworld ( perhaps one reserve entrance)
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