Glukkons' Legs
I'm surprised that nobody has mentioned that the Glukkons once had legs that they walked on. Executive worker class Glukkons walk on their hands because their legs are only "vestigial" stubs. Since vestigial is defined as a remnant of something that had once existed, it is apparent that the Glukkons once had four limbs rather than the two that you see in the past Oddworld games. You can see the tiny little "legs" hanging from Molluck in Abe's Oddysee's good ending when his suit is electrocuted off. (They're not what he was walking on.)
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