What?? You talkin' to me??
*removes earphones*
Reading through your speculation, I found myself engrossed in the storyline. It's not often I find a good fan theory that does that. That alone has to be congratulated, so here goes...
Congratulations.
Species X sounds like it could be a very interesting race to meet sometime in the future. From the story you've given, I doubt that they would have grown up to be at the top of the food chain, it sounds to me rather like they're either extinct or have dwindled down to somethat that is hardly known about in Mudos. I love the idea of the Mudokons having a darker history than any of them could today imagine, and I can imagine Abe finding out and deciding what to do next, while the other heroes are all around him feeling kind of guilty or whatever. I would like to see this Species X, and I like the idea of Abe visiting them someday.
Now, with a slight alteration to your story, I can see how Species X could easily have developed into the Khanzumerz we know [of] today. Perhaps they didn't lose the battle with the Glukkons after all, perhaps they were going to win, but something made them change their mind. They sound like a rather shallow-minded people (just like I imagine the Khanzumerz), and I can see how they could have been tricked by the Glukkons to step down their attacks. But I think that it's more likely Species X turned against the Mudokons for some reason (feel free to speculate), and since they were the dominant race in Mudos (empire-wise), decided that the best way to upset the Mudokons was by giving the Glukkons a helping hand. In fact, I can see the Khanzumerz being the real catalyst for Mudos's Industrial Era, but perhaps not by desiring consumer goods, originally.
This would make Khanzumerz not only the most important family in Mudos in terms of Moolah, it would also mean they've got some kind of claim over the lands or the politics. While Mudos doesn't seem to have any governing body of sorts, this dominance would explain how the Khanzumerz have come to earn such a respect from their industrial friends. Perhaps the Khanzumerz are, to this day, a warlike species, although I don't think so. I do think they still have a grudge against the Mudokons, which would be why they are truely evil, and perhaps they still remember the days when they were 'natives', and want to put an end to all things native and natural in Mudos.
Well, that's just a few thoughts made up as I type along.
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