[One, Two, Middlesboogie]

Evolution in the truest sense of the word is the gradual change of a species's anatomy in order to suit its environment, determined by natural selection and often resulting in the creation of a new species.

Since Glukkons were obviously designed with Earthly squids and cuttlefish in mind, I think Glukkons and the other octigians evolved from water creatures. I think the Glukkons' ears must've originally been propulsion siphons like Earthly octopi... rather in the same way as our inner ear bones evolved from reptilian jaw hinges.

Although, your post seemed to be more about history rather than biology. The Magog Cartel was originally created with good intentions; to provide jobs for the Mudokons. But it went further and further downhill for some reason, and the Glukkons becae nastier and nastier, and their reputation went out the window as well.