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So this spiteful dead souls locked in bottles (...) became something very horrible and unpleasant, finally got escape and bring their wrath down upon the first living beings they see - the innocent Muds. Ghosts eating their souls from inside.
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Wait a minute, is the Age of Alchemy still a thing in Glukkon history?
Because now that I think about it, Soulstorm brew sounds like a fusion between an addictive soft drink and literal potion/curse making, what with the bones and tears being used as ingredients.
EDIT:
And I'm pretty sure the Three Weirdos flat-out confirm that the bones are tied to the ghosts, considering they're missing limbs because their bones got used for brew.