AE was never that dark to begin with except for thematically. NNT was literally taking AE's tone and slapping it onto AO and all the discussions I had with Sock at the time were saying "I sure don't think AE's tone fits AO at all even if that's the way Lorne always wanted Oddworld to be". AE has always been a more lighthearted and goofy game comparitively, it set the stage for the mood of Oddworld from that point onwards even if MO over did the goofy aspect of it, and NNT eventually made it to the point where Lorne wanted to make it his "original vision" of Oddworld which just so happened to have the same tone as AE.
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