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I'm still not entirely sure how NnT is that much more comedic than Oddysee or Exoddus, just because it has more expressive characters and a couple of silly one liners from the Mudokons.
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Mudokons are constantly spouting one-liners and portrayed as oafish and upbeat. Sligs are no longer menacingly silent but are constantly muttering away about their paycheck and how much they wanna shoot stuff with their guns. Abe lets out a buffoonish laugh every time he kills a Slig.
The characters aren’t more expressive, they’re just more exaggerated.