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I'm more just saying it's a sense of consistency. since they seemd to have locked down a tone in AE (which isn't necessarily lighter - since AE's plot is pretty dark) which is probably more the fact that the characters (Mudokons, Sligs, Abe) have more humourous elements to them. So it makes sense to add these to a remake of AO since there's a consistency in the characters and less of a noticeable change between NnT and future installments of the Quintology. But besides from a couple of moments in the cutscenes from I've seen the majority of this takes place within gameplay and the dialogue heard by Mudokons and Sligs. And, I mean, they even took the most cartoonish cutscene from AO and made it relatively a lot darker - so they haven't abandoned that side of the storytelling as well.
But yeah that'll be my final word too I guess. unless Manco replies with a harshly worded comment....
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The characters were consistent as they were and showed visible progress with each game, even in MO. There would be no visible or clear character development if you portrayed them exactly the same every appearance they had.