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Really? Essentially every cutscene in AE was full of jokes or humour. It was dark and grim as well but I don't see how NnT is much more humourous than AE to the point of going past MO's more cartoonish feel. Only the ingame characters are more humourous and to me that's just the characters in game lining up to the characters you see in the cutscenes.
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A lot of AE’s humor is quite deadpan in its delivery, which helps with its integration into the larger world presented in the game. Take the MOM interviews with Phlegm, Dripik and Aslik – all three are portrayed as incompetent in some way, but it’s entirely through what they’re saying. The player sees this incompetence but nobody calls them on it in the game and the joke isn’t lingered on. Wordplay like this doesn’t come across as aggressively cartoonish and unrealistic as visual slapstick like a character tripping and falling while accompanied by goofy sound effects.
As to “characters in game lining up to the characters you see in the cutscenes”, this just further takes away from the atmosphere as it spreads humor out through the entire game, making it more common and therefore allowing the player to grow accustomed to it. It doesn’t allow a contrast to develop between the dark and the comedic, as you are constantly being presented with both.