MOD EDIT: Split from this thread.
Quark was a/the villain in the first and second games. After that he's been a bumbling support character.
This is going to be awful. I have very low expectations. Story has never been the strong suit of Ratchet and Clank. The first 3 games barely took themselves seriously, which worked well. The biggest downfall of the Future trilogy was the serious and poorly written story. I'm not even going to mention "All 4 One!".