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Neither. It uses some kind of PSX emulator to emulate system calls, graphics etc., but it runs native x86 code. Sort of like compiling Windows app to Linux with Winelib.
AO and AE use MIDI for all music, but it doesn't use the crappy gm.dls that comes with Windows but custom soundfonts created for the game (They're a part of sounds.dat). MIDI is, in fact, a very good format as long as you use good soundfonts.
Also, can someone explain what's wrong with the PC port of AO/AE? I find it pretty accurate and well made...
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I agree. The PC ports are graphically and technically superior to the PSX versions. The MIDI music is the same as what runs on the PSX too (SEQ is midi with a soundfont).
I only find that the music sounds better on PSX due to the PC SEQ player being buggy.