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I don't know if anyone's mentioned it, but I always thought it was kind of -odd- that you began rescuing Mudokons in the first level. Who told Abe he could chant to open portals? That seems like something he'd have picked up in his travels through Monsaic Lines. I always felt like the escape from Rupturefarms should have been dedicated to learning the controls and that returning to Rupturefarms to rescue Mudokons should have been the reason Abe was brought to Monsaic Lines in the first place.
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I like having Mudokons there from the start. It adds a little more depth and realism to the world—would be rather strange having one single area of RuptureFarms begin completely devoid of Mudokons—and it creates a little replay value for players who missed them first time and want to go back for a perfect run.
I also think that the controls for Abe are fairly simple to pick up, but the GameSpeak needs a fair bit of introduction at the beginning. So having Mudokons around helps players in that regard.