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Maybe there could be Ratz in some areas every now and then. They are closely connected to the Raisin though, so I'm not sure whether there should be a reference to him that early in the story.
Another thing I was thinking about: the bell songs. In AO, Abe would always do them for you when you got them from the story stones. I think it would be neat if you had the choice to memorize them or have Abe memorize them for you.
Also, I don't think co-op would hurt if it was a completely separate story with no real connection to the Quintology. I like the game mechanics of AO and AE and would love to see them used in a co-op game. I wouldn't want to see the players rescuing mudokons through bird portals or anything, though. The role of the savior is best left for Abe, I think.
If it's going to take place in RuptureFarms, then I think it shouldn't be the one Abe was working at. The connections to the other games should be loose. I feel that Oddworld doesn't need to be a single player experience, but for the most part it should be. Especially the Quintology.
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I think co-op could work in the same story as single player if it was the same kind of single player in AO, but had support for more players, LAN, and online.
There could also be multiple game modes that are separate from the story. Like a siege mode, where players take turns defending and attacking a factory with a flag in it. The defensive team could be the industrialists and the offensive team can be the Mudokons.