I thought they did, but wasn't too sure on clarification. Shoulda checked really.
I'd take it that their queen system evolved independently from the other species, in this case, after said split off from the lineage that lead to Scrabs, and possibly Meeches (Paramites and Scrabs retained the original leg structure, where the Meech went on to change, also showing how they can be similar to, say, Elum but are on a different angle somewhat).
This may also show that the Glukkon lineage is more separated from the others and may also be convergent queen system. Maybe they are all separate, since it appears to work well on Oddworld, thus why it has cropped up often.
Either that or Shrykull did something. This god, at least, does physically exist within the universe. Or perhaps, the Shrykull spirit was created in respect of Scrabs and Paramites by a number highly powerful shamans from the Mudokons foggy prehistory? (either from scratch or with a representative from each species) Hence it wouldn't exactly have godly power over species evolution.
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