You should have looked at wheat. There are no clades smaller that
Triticum because they hybridise so often. Cereal sluts!
Genuine hybrid species are rare in the animal Kingdom, because polyploidism, which is required for a hybrid to become fertile, is usually deleterious in animals. I don't know how Gloktigi work, but possibly they're more like mules.
Species with queens present no particular difficulty to cladistics.
EDIT: In the very thread you linked to I find this awkward acknowledgement:
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I'm loathe to admit the existence of either [magic or psychic abilities], but since this is a fictional world in which several characters appear able to perform amazing world-altering feats by some unseen and seen power, we must entertain the possibility of there being fantastic abilities accessible to mudokons at the very least.
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Which I must use to allow any weird and impossible evolutionary relationships, obscene as they may appear to me. It does render discussion sans officially released palaeobiology rather futile, however.