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completely underwater, where the atmosphere is dark and misty, and there are creatures moving in and out of the dark, including a giant Sea Rex slithering its way through the background, yet somehow remains the focus of attention, considering the player shouldn't really know what they're supposed to be doing yet. I think a level like that, with the whole game following the same extremely creepy suit, would be amazing.
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There is a ton of things that could be done in an underwater level. It could be like a free-for-all; no specific paths to take, and lost of little tunnels to explore in. Plus there could be islands and boats and stuff to hop up on that could make things much more interesting.
Thinking of underwater levels made me think of the 'dark zone', that empty void at the bottom of the ocean where everything is pitch black. A lantern-fish oddworld creature would be cool beyond comparison.