I.E. starts up quicker to my knowledge while Firefox once loaded runs a hell of a lot faster due to better caching capabilities i guess.
That said the start up on firefox is only about 5 or so seconds longer then I.E.
All in all the popularity of Firefox is probably more due to its add on capabilities and user community.
Any body who wants to and knows how can contribute to the mozilla community if they wish which leads to a large amount of useful tidbits that can be added to firefox for optimal browsing.
For instance i dont even have to go to photobucket to upload images to my account.
The same applies to my blog.
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