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You did see the canyons and chasms, right?
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Canyons don't mean the river was especially big. For the Mongo to be as big as it is 'required' to be, it would need to be bigger than pretty much any river we have on Earth. I don't think there was *quite* the scale of the grand Canyon. All those mean is essentially how long the river's been there to erode the terrain like that.
I haven't played the game, but I'm judging from what I've seen here. I just accept the fact that either Ma'Spa is as big as an entire town or something, or that there are other equivalents to it, otherwise I just can't see how that would work for supporting such a populous species.
I'm fairly sure Thudslugs can feel 'love', if that's the way you want to put it, otherwise they wouldn't breed and soon become extinct.