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When did we ever hear that they go to Ma'Spa to develop lungs?
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‘
Gabbiwogs develop lungs after a dip in the spawnin springs.’
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That might refer that Ma'Spa is an ancient city. And Munch's biography said that he just came from a spawning pool high up in the mountains, so it's very well possible that it isn't Ma'Spa.
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It’s very possible that Gabbits come from other rivers as well as the Mongo, but no reason to believe they do. Ma’Spa is a breeding ground, and has been sculpted into a shrine, but what makes you think there’d be a city there?
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Why does he live now in a cavern? And how did Abe meet him? And what are those mountains mentioned in the biography?
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He has been driven underground to hide from the industrialists. As a figurehead for native and environmental rights, they would love to have him out of the picture. As a tree that’s developed into a thinking, talking creature, the Vykkers would love to tear him to pieces.
Munch’s Oddysee is meant to take place in a mountainous area. It doesn’t really show.