It has not been released. There are several theories, including:
1. That the tail controls the legs.
2. That the wires to the mask somehow send messages from the head.
3. That the legs have sensors that can see pulses in the nerves in the slig-bottom.
Do a search and you'll find all the discussions about them.
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Spending as long as I do here, it's easy to forget that Oddworld has actual fans.
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