My personal idea, totally unfounded except by just..practical purposes, is that Abe has no nose whatsoever. Look at the illustrations of mudokon skulls. There are no nostrils or any indication of something on the skull itself which would perform that function.
I always thought mudokons would have something similar to the Jacobson's organ of a cat. This is an organ stimulated during the process of flehming, which I'm sure most cat-owners have seen their pet do. When it smells something new, the cat opens its mouth and passes air over the organ for prolonged periods, so as to better categorize the properties of a scent. Since mudokons can CLEARY smell, what with going "phooh!" when they smell Abe farting, there has to be some method of sensing odors. Something that could be "smelled" as air passes into the mouth seems logical and uncomplicated to me.
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