Now that I think about, maybe when the Glukkons were enslaving Mudokons the temple was connected to the ground. Yet when the Glukkons captured Scrabs, the Scrabs got scared and hid in the temple for protection. The Mudokons probably tried to protect the Scrabs from the Glukkons, yet the Glukkons probably cut off all ground enterances for the Mudokons to get into the temple from. Maybe some Mudokons used the wells to get into the temple from secretly. Yet too many Industerial possibilities are open with this idea.
To respond to Zerox's question, there is plenty of small animals in the temple. There are flocks of bats that Scrabs probably use for food. There are also birds that Scrabs could sneak up on, or maybe the Slogs and Sligs that invade the temple could be used for food.
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