Never seen Abe keep his mouth open long enough for breathing, nor doing anything with his tongue.
Maybe that feathery thing on his head absorbs gases from the atmoosphere into a canal that goes down the neck to lungs. That's why he still chokes and grasps his neck in that killer gas stuff in the PS1 Oddworld games, and absorbing gases would mean he'd sense a smell at the same time.
But an actual nose like birds would make more sense...maybe hidden in those creases above his mouth we always see. They are there.
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