Molluck probably didn't have them because Gloktigi hadn't been introduced into the Oddworld universe yet or thought of.
Maybe Molluck wasn't quite important enough for that protection...the only others we know are Lady Margeret herself and Sekto, who owned a whole river, which I'm sure is worth more than Rupturefarms.
I can't remember where the Mudokon/bird idea came from/ But thinking more about it...that would explain the hand/foot structure quite well, if we look at bird skeletons.
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