I think that you got it all wrong. Abe and Munch are not on the bottom of the food chain, but in the middle. They are referred to as weak characters, since the enemies they face either are powerful hunters or people that kill them for sport. Weak doesn't mean 'bottom of the food chain'.
OK, I will explain the food chain concept. On a food chain, there are animals (humans are animals too you know) that are connected to each other by their consuming related activities. When an animal is lower on the food chain than they are of another one, they are considered prey if they are connected. That means the one higher requires them as food. Now let's look at Mudokon food chain, hmm? Pretty much every Mudokon requires some meat, so they are in the middle, since they too are hunted. Gabbits' are similar. They feast on Worryfish, but are hunted down by Searexes. So none of them are on the bottom.
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