Where they? It was just mollusc-like descendants to my knowledge, explaining the head. That doesn't mean they lived in swamps.
But...that raises another question. How did Glukkons lose their legs? For vestigial legs to be there, usable elgs must have been there before. So why did the Glukkons lose their legs?
If you're talking about Glukkons and their current build, what is known is that they once were at peace with the Mudokon tribes. I presume Glukkons formed vaguely similar tribes of a sort on plain, flat ground. The long arms and all would be well suited to walking quickly across such an environment. Except now they were very restrictive suits. For some reason. Morons.
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