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If they're based on beetles, then they have a hard shell as an exoskeleton but no internal bones.
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Oh dear. We'd better ignore Clakkers. Birds with breasts, indeed. And lizards with feathers. Who are these Mudokons. Oddworld's species evolved on Oddworld, they're not bound by the same taxonomy as Earth creatures. The beetle at the start of
Munch's Oddysee strongly suggests Meetles have skeletons, which they would need to support their size anyway.
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Lorne has created a great universe, however, I think it's the fans who make the biggest contributions. … The gap between Glukkons and Octigi for instance. Lorne only said they were related somehow, but it wouldn't surprise me if he didn't even exactly make up how himself yet.
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Well, we wouldn't be a very good audience if we didn't use our imagination to fill in the gaps.
Besides, we know Glukkons and Octigi are related because they both belong to the same taxon along with Gloktigi.