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I believe the longevity of the game is going to fall down to creativity in evolving creatures and designing their buildings. Apart from that, I imagine the gameplay gets stale quickly, and even the novelty of that's going to drain, especially for the kids who take things to extremes by adding just those few hundred extra legs.
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I wouldn't be surprised if the gameplay did get quite stale. Some other features to make the game more enjoyable and playable would be quite cool and, I think, neccessary.
However, creating and evolving my own species would be incredibly cool and interesting, and that alone would keep me entertained for tens of hours.