Theories include:
Colour depends on spirituality
Colour in chameleonic through other ways, such as enviroment
Colour depends on mood
Colour depends on father/genes
Colour depends on exposure to pigment-changing things, such as UV rays
Colour depends on skin-paint.
AE was the game that basically confirmed the mood idea, even though the AE mudokons only changed colour for incredibly brief periods of time.
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