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Ah, I'm not the only one to remember that. Trouble is, I can't remember whether it was actually on Oddworld.com, or just in a fan theory. I can remember a fan theory on a website that distinctly elaborated on what I knew was on Oddworld.com, and the History text from the AE-era Oddworld.com makes no mention of Glukkons being mutated by the disaster at the end of their Alchemy Era.
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Well if Glukkons have unusable legs, that would imply that at some point in evolution they looked completely diffrent (stating the obvious).
Could it perhaps be that (since Glukkons, Octigi and Glocktigi are related) that Glukkons are the mutated version of Octigi or Glocktigi. This would somewhat explain their unused legs and their oversized heads.
Just like Octigi, Glukkons don't realy have a body so it's rather reasonable to asume they are mutated from the Octigi.