If Steef are the ancient protectors of the Grubbs and the river then killing them serves the practical purpose of destroying the defenders of the river you want to own. Once Sekto rose to power he started hunting the Steef in order to quell opposition.
Also there's the possibility of a prophecy concerning Steef vanquishing Octigi. Thats what the rock art in the mongo valley seemed to imply.
I'm assuming Steef heads became valuable because Sekto was collecting them. Who knows when the idea of Steefs as freaks came into vogue? We know the Clakkerz couldn't have been there before the dam, and it makes sense that the outlaws followed the clakkerz. So I'm thinking it all starts with Sekto. But yeah, it could be a vengeance thing for him.
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