I think it has always been established that Scrabs and Paramites obey the muds because they treat them with respect. The Scrabs and Paramites that abe experiences, are creatures that have adapted to become aggressive toward everyone because of the glukkons.
Paramites probably aren't nocturnal, just more active during the night, and fairly sluggish/inactive during the day. I don't doubt that the paramites of AO and AE live in the darkness because it's safer for them.
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