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I rather Scrabs and Paramites didn't have a sense of spirituality, and were simply wild creatures. I don't think they did service to the Mudokons by protecting them, but the Mudokons in fact just took advantage of their voracious temperaments to keep dangers from entering their temples or vaults. Maybe the Guardians were just wild creatures, too, and are just like the Meeches in that their legacy has been virtually completely forgotten.
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I understand where your coming from, but how do you explain the Scrabs and Paramites standing on top of the "totems" in the Monsaic Sanctum? If they were wild, I'd expect the Paramites to run off, from fear of the massive crowd of Mudokons, and I'd expect the Scrabs to start fighting ith eachother, or turn to the Mudokons in a bid for a quick feed.
Something is obviously keeping them up there. I believe it's a sense of spiritualism. Or maybe it's the forces that be
Alcar...